We selected nine study sites, each incorporating three vegetation states: (a) fallow cropland, representing the restoration starting point, (b) planted old field (actively restored site), and (c) reference York gum (E. loxophleba) woodland.

Plant species richness and cover

All annual and perennial plant species were recorded in spring 2017 within each plot and identified to genus and species level where possible. Nomenclatures follow the Western Australian Herbarium (2017). A point intercept method previously demonstrated to provide objective and repeatable measures of cover (Godínez-Alvarez, Herrick, Mattocks, Toledo & Van Zee 2009; Prober, Standish & Wiehl 2011) was used to quantify cover of individual plant species, total vegetation cover and substrate types (i.e., bare ground, litter cover, plant cover). Ground cover, individual species, and canopy cover intercepting at every 2 m along four parallel, evenly spaced 50 m transects across each plot were recorded using a vertically placed dowel (8 mm wide, 2 m tall), resulting in 100 intercepting points per plot. For planted old fields, transects were placed parallel to planting rows, with two centred on rows and two centred between rows. This approximately represented the relative abundance of planted rows and non-planted inter-rows. If a species was recorded in the plot but did not intercept the dowel on any transect it was assigned 0.5 points. This method provided a measure of relative abundance (percentage cover) of plant species across the plot.

To calculate species richness and cover across different life history and growth forms, species were classified into the following groups: total, native trees, native shrubs, native non – planted shrubs, native grasses, native perennial forbs, native annual forbs, exotic grasses and exotic annual forbs using the Western Australian Herbarium (2017) classification.

Woody debris and leaf litter surveys

Leaf-litter dry mass was estimated by collecting leaf-litter from five randomly placed 25 cm x 25 cm quadrats along two 50 m transects across each plot. Litter was stored in paper bags for transportation and then oven dried for 36 hours at 60 °C. The dried litter was weighed to 3 decimal points. Cover of fine and coarse woody debris and litter depth was estimated at every meter along two 20 m transects for each plot. Woody debris was classified by diameter. Length, max and min diameter was measured for all logs with a diameter greater than 10 cm.